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Frank, P. W.: Growth rates and longevity of some gastropod mollusks on the coral reef at Heron Island. Oecologia 2, 232–250 (1969)

Connell, J. H.: The influence of interspecific competition and other factors on the distribution of the barnccle Chthamalus stellatus. Ecology 42, 710–723 (1961)

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Anonymous Tide Tables: High and low water predictions. West Coast of North and South America, including the Hawaiian Islands. U.S. Dept. Commerce, National Oceanic and Atmospheric Administration (1963–1973)

Along exposed rocky intertidal shorelines of western North America the mussel Mytilus californianus exists as a characteristic, well-defined band. Measurements at Mukkaw Bay and Tatoosh Island, Washington State, suggest that the upper limit to distribution is constant. The lower limit is also predictably constant, as judged by photographs of the same areas taken up to 9 years apart. The band of mussels is formed by larval recruitment to a variety of substrates, especially the filamentous red alga Endocladia muricata. From the settlement site, if the mussels survive a series of predators including the starfish Pisaster ochraceus and a variety of carnivorous gastropods (Thais spp.), the mussles may be washed inward or migrate (be pushed) downward.

When Pisaster was removed manually, the zonation pattern changed rapidly. Mussels advanced downward at Mukkaw Bay a vertical distance of 0.85 m in 5 years. No movement was observed on 2 adjacent control sites. At Tatoosh Island a maximum displacement of 1.93 m has been observed in 3 years; the slope there is 40°. Again, there was no change at control sites with Pisaster. At Mukkaw Bay over 25 species of invertebrates and benthic algae are excluded from occupancy of the primary substratum by mussels. The ecological dominance of mussels is discussed; predation is shown to enhance coexistence among potential competitors. A survival curve for Pollicipes polymerus indicates that the time course for interspecific competitive exclusion may be long (76 months). The clarity of the biological interrelationships and the constancy of pattern through time provide no support for the contention that intertidal communities are physically-controlled.

Glynn, P. W.: Community composition, structure, and interrelationships in the marine intertidal Endocladia muricata-Balanus glandula association in Monterey Bay, California. Beaufortia 12, 1–198 (1965)

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Feder, H. M.: Growth and predation by the ochre sea star, Pisaster ochraceus (Brandt), in Monterey Bay, California. Ophelia 8, 161–185 (1970)

Paine, R. T.: The Pisaster-Tegula interaction: prey patches, predator food preference and intertidal community structure. Ecology 50, 950–961 (1969)

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Menge, B. A.: Competition for food between two intertidal starfish species and its effect on body size and feeding. Ecology 53, 635–644 (1972)

Paine, R. T., Vadas, R. L.: The effects of grazing by sea urchins, Strongglocentrotus spp., on benthic algal populations. Limn. Ocean. 14, 710–719 (1969)

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Chan, G. L.: Subtidal mussel beds in Baja California with a new record size for Mytilus californianus. Veliger 16, 239–240 (1973)

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Luckens, P. A.: Breeding, settlement and survival of barnacles at artifically modified shore levels at Leigh, New Zealand. N.Z.J. mar. Freshwat. Res. 4, 497–514 (1970)

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Dayton, P. K.: Dispersion, dispersal and persistence of the annual intertidal alga, Postelsia palmaeformis Ruprecht. Ecology 54, 433–438 (1973)

Batham, E. J.: Pollicipes spinosus Quoy and Gaimard. I. Notes on the biology and anatomy of adult barnacle. Trans. roy. Soc. N.Z. 74, 359–374 (1945)

Ricketts, E. F., Calvin, J., Hedgpeth, J. W.: Between pacific tides. Stanford, California: Stanford University Press 1968

Seed, R.: The ecology of Mytilus edulis L. (Lamellibranchiata) on exposed rocky shores. I. Breeding and settlement. Oecologia 3, 277–316 (1969)

Hewatt, W. G.: Ecological succession in the Mytilus californianus habitat, as observed in Monteray Bay, California. Ecology 16, 244–251 (1935)

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Landenberger, D. E.: Studies on selective feeding in the Pacific starfish Pisaster in southern California. Ecology 49, 1062–1075 (1968)

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Wilson, D. P.: Some problems in larval ecology related to the localized distribution of bottom animals, p. 87–99. In: Perspectives in marine biology, A. A. Buzzati-Traverso, Ed. Berkeley: University California Press 1958

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Paine, R. T.: A short-term experimental investigation of resource partitioning in a New Zealand rocky intertidal habitat. Ecology 52, 1096–1106 (1971)

Transformers, extension leads and connectors are all available under the ‘Click & Build’ section of our website to allow you to truly customize your lighting project.

Connell, J. H.: A predator-prey system in the marine intertidal region. I. Balanus glandula and several predatory species of Thais. Ecol. Monogr. 40, 49–78 (1970)

Dayton, P. K.: Competition, disturbance, and community organization: the provision and subsequent utilization of space in a rocky intertidal community. Ecol. Monogr. 41, 351–389 (1971)

Hatton, H.: Essais de bionomie explicative sur quelques especes intercotidales d'algues et d'animaux. Ann. Inst. Oceanogr. Monaco 17, 241–348 (1938)

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Harger, J. R.: Competitive co-existence: maintenance of interacting associations of the sea mussles Mytilus edulis and Mytilus californianus. Veliger 14, 387–410 (1972)

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Paine, R.T. Intertidal community structure. Oecologia 15, 93–120 (1974). https://doi.org/10.1007/BF00345739